Anurans of In Becetèn (Republic of Niger): the most diverse site for amphibians in Mesozoic Africa

In Becetèn (Niger) represents one of the most diverse continental vertebrate faunas from the Late Cretaceous of Africa. Recently, anurans, in particular aquatic pipimorphs, have generated renewed interest, with the identification of four distinct pipimorph taxa. Here we describe the remaining anuran specimens from In Becetèn, which represent at least three new anuran taxa that cannot be assigned to pipimorphs (Anura indet, Neobatrachia and a ranoid). Among them, one taxon, documented by ornamented cranial material, is a large Neobatrachia incertae sedis, as suggested by our phylogenetic analyses. This marks only the third Mesozoic occurrence of neobatrachians in Africa. In addition, an isolated humerus is referred as a new unnamed ranoid and is not only the oldest known occurrence of the clade, but also the first known Mesozoic specimen of this cosmopolitan family from Africa. With at least seven anuran taxa identified, In Becetèn is the most diverse locality of Africa, and one of the most diverse localities across the Late Cretaceous (Coniacian or Santonian).