Integrative taxonomy supports the description of two new species of Macrobiotidae (Tardigrada, Eutardigrada) from Kristianstads Vattenrike Biosphere Reserve in Sweden

Kristianstads Vattenrike Biosphere Reserve (KVBR) in the south of Sweden is a known hotspot for tardigrades, with almost a third of the country’s tardigrades species reported from this location alone. Herein, the tardigrade diversity of the KVBR is further investigated. New records for the country are reported and two new species, Macrobiotus vattenrikense sp. nov. and Paramacrobiotus marchelmoni sp. nov., described using an integrative taxonomic approach. An updated multi-locus phylogeny to the family Macrobiotidae is provided with the inclusion of the newly described species.