New records of Frullania Raddi (Frullaniaceae, Marchantiophyta) from Peru including notes on F. tetraptera Nees & Mont. and its synonyms

Nine Frullania Raddi species are reported new for Peru, belonging to F. subg. Chonanthelia Spruce (F. beauverdii Steph., F. cuencensis Taylor, F. holostipula S.Hatt. & D.G.Griffin, F. pluricarinata Gottsche, F. winteri Steph.), F. subg. Homotrophanta Spruce (F. nodulosa (Reinw., Blume & Nees) Nees), F. subg. Meteoriopsis Spruce (F. intumescens (Lehm. & Linden.) Lehm. & Lindenb. var. intumescens, F. lobatohastata Steph.) and one incertae sedis species (F. pyricalycina Steph.). Androecia are newly described for F. beauverdii Steph., a species previously considered endemic to Bolivia. Original material of F. tetraptera Nees & Mont. as well as additional Peruvian specimens of this species are dioicous and deviate from earlier published circumscriptions. Frullania conferta Spruce, formerly considered a synonym of F. tetraptera Nees & Mont., is reinstated as a distinct species based primarily on differences in its monoicous sexual condition as well as lobule, styli and gynoecial bract morphology. Another prior synonym of F. tetraptera, F. semiconnata Lindenb. & Gottsche, is newly proposed as a synonym of F. mexicana Lindenb. Frullania mexicana differs from F. tetraptera and F. conferta in its gynoecial bract morphology and number of perianth keels. Lectotypes are designated for F. conferta, F. cuencensis, F. semiconnata and F. tetraptera.